The paternal relationship between Waverly Oren Upton and his daughter Frances Vivian Upton is confirmed by the direct inheritance path of autosomal DNA down to their DNA tested descendant SRB.

The segmental (via DNA segment triangulation) and total DNA shared between SRB and other confirmed descendants of Charles Perry Upton/Celinda Ellen Gray is consistent with the expected ranges for their respective relationships. Since SRB descends from Charles Perry Upton/Celinda Ellen Gray through Waverly Oren Upton and his daughter Frances Vivian Upton, the presence of significant shared DNA between SRB and other known descendants of Charles Perry Upton/Celinda Ellen Gray validates the generational links along this paternal line.

This supports the conclusion that DNA was passed from MRCA couple Charles Perry Upton/Celinda Ellen Gray through Waverly Oren Upton to Frances Vivian Upton, and ultimately to SRB, confirming this direct ancestral connection through multiple generations.